STRUGGLE FOR EXISTENCE BEARS ON NATURAL SELECTION GEOFFROY SAINT HILAIRE IN ZOONOMIA , DR. W.G. WELLS 1813 , PATRICK MATTHEW IN NAVAL TIMBER AND ARBORICULTURE, PROFESSOR OWEN IN 1849, PROFESSOR HUXLEY 1859, Dr. HOOKER 1859 And So On STIFINS HAWKINS; WHAT VEDAS OF VEDIC TRADITIONS HAVE ALREADY ESTABLISHED

To be sure, the contradictions between evolution and the Bible have been addressed again and again. In fact, the critical issues were successfully resolved long before Darwin published his views on natural selection in 1859. What has not been repaired is the breach that opened up between science and religion. The struggle for existence bears on Natural Selection. Geoffroy Saint-Hilaire's ('Hist. Nat. Générale,' tom. ii. p. 405, 1859) excellent history of opinion on this subject. In this work a full account is given of Buffon's conclusions on the same subject. It is curious how largely my grandfather, Dr Erasmus Darwin, anticipated the views and erroneous grounds of opinion of Lamarck in his 'Zoonomia' (vol. i. pp. 500-510), published in 1794. According to Isid. Geoffroy there is no doubt that Goethe was an extreme partisan of similar views, as shown in the Introduction to a work written in 1794 and 1795, but not published till long afterwards: he has pointedly remarked ('Goethe als Naturforscher,' von Dr Karl Medinge s. 34) that the future question for naturalists will be how, for instance, cattle got their horns, and not for what they are used. It is rather a singular instance of the manner in which similar views arise at about the same time, that Goethe in Germany, Dr Darwin in England, and Geoffroy Saint-Hilaire (as we shall immediately see) in France; came to the same conclusion on the origin of species, in the years 1794-5.
Geoffroy Saint-Hilaire, as is stated in his 'Life,' written by his son, suspected, as early as 1795, that what we call species are various degenerations of the same type. It was not until 1828 that he published his conviction that the same forms have not been perpetuated since the origin of all things. Geoffroy seems to have relied chiefly on the conditions of life, or the 'monde ambiant' as the cause of change. He was cautious in drawing conclusions, and did not believe that existing species are now undergoing modification; and, as his son adds, "C'est donc un problème à réserver entièrement à l'avenir, supposé meme que l'avenir doive avoir prise sur lui.'
In 1813, Dr W. C. Wells read before the Royal Society 'An Account of a White female, part of whose skin resembled that of a Negro'; but his paper was not published until his famous 'Two Essays upon Dew and Single Vision' appeared in 1818. In this paper he distinctly recognises the principle of natural selection, and this is the first recognition which has been indicated; but he applies it only to the races of man, and to certain characters alone. After remarking that negroes and mulattoes enjoy an immunity from certain tropical diseases, he observes, firstly, that all animals tend to vary in some degree, and, secondly, that agriculturists improve their domesticated animals by selection; and then, he adds, but what is done in this latter case 'by art, seems to be done with equal efficacy, though more slowly, by nature, in the formation of varieties of mankind, fitted for the country which they inhabit. Of the accidental varieties of man, which would occur among the first few and scattered inhabitants of the middle regions of Africa, some one would be better fitted than the others to bear the diseases of the country. This race would consequently multiply, while the others would decrease; not only from their inability to sustain the attacks of disease, but from their incapacity of contending with their more vigorous neighbours. The colour of this vigorous race I take for granted, from what has been already said, would be dark. But the same disposition to form varieties still existing, a darker and a darker race would in the course of time occur: and as the darkest would be the best fitted for the climate, this would at length become the most prevalent; if not the only race, in the particular country in which it had originated.' He then extends these same views to the white inhabitants of colder climates. I am indebted to Mr Rowley, of the United States, for having called my attention, through Mr Brace, to the above passage in Dr Wells' work.
The Hon. and Rev. W. Herbert, afterwards Dean of Manchester, in the fourth volume of the 'Horticultural Transactions,' 1822, and in his work on the 'Amaryllidaceae' (1837, pp. 19, 339), declares that 'horticultural experiments have established, beyond the possibility of refutation, that botanical species are only a higher and more permanent class of varieties.' He extends the same view to animals. The Dean believes that single species of each genus were created in an originally highly plastic condition, and that these have produced, chiefly by intercrossing, but likewise by variation, all our existing species.
In 1826 Professor Grant, in the concluding paragraph in his well-known paper ('Edinburgh philosophical journal,' vol. xiv. p. 283) on the Spongilla, clearly declares his belief that species are descended from other species, and that they become improved in the course of modification. This same view was given in his 55th Lecture, published in the 'Lancet' in 1834.
In 1831 Mr Patrick Matthew published his work on 'Naval Timber and Arboriculture,' in which he gives precisely the same view on the origin of species as that (presently to be alluded to) propounded by Mr Wallace and myself in the 'Linnean journal,' and as that enlarged in the present volume. Unfortunately the view was given by Mr Matthew very briefly in scattered passages in an Appendix to a work on a different subject, so that it remained unnoticed until Mr Matthew himself drew attention to it in the 'Gardener's Chronicle,' on April 7th, 1860. The differences of Mr Matthew's view from mine are not of much importance; he seems to consider that the world was nearly depopulated at successive periods, and then re-stocked; and he gives as an alternative, that new forms may be generated ' without the presence of any mould or germ of former aggregates.' I am not sure that I understand some passages; but it seems that he attributes much influence to the direct action of the conditions of life. He clearly saw, however, the full force of the principle of natural selection.
The celebrated geologist and naturalist, Von Buch, in his excellent 'Description physique des Isles Canaries' (1836, p. 147), clearly expresses his belief that varieties slowly become changed into permanent species, which are no longer capable of intercrossing.
Rafinesque, in his 'New Flora of North America,' published in 1836, wrote (p. 6) as follows:- 'All species might have been varieties once, and many varieties are gradually becoming species by assuming constant and peculiar characters'; but farther on (p. 18) he adds, 'except the original types or ancestors of the genus.'
In 1843-44 Professor Haldeman ('Boston journal of Nat. Hist. U. States, vol. iv. p. 468) has ably given the arguments for and against the hypothesis of the development and modification of species: he seems to lean towards the side of change.
The 'Vestiges of Creation' appeared in 1844. In the tenth and much improved edition (1853) the anonymous author says (p. 155):- 'The proposition determined on after much consideration is, that the several series of animated beings, from the simplest and oldest up to the highest and most recent, are, under the providence of God, the results, first, of an impulse which has been imparted to the forms of life, advancing them, in definite times, by generation, through grades of organisation terminating in the highest dicotyledons- and vertebrata, these grades being few in number, and generally marked by intervals of organic character, which we find to be a practical difficulty in ascertaining affinities; second, of another impulse connected with the vital forces, tending, in the course of generations, to modify organic structures in accordance with external circumstances, as food, the nature of the habitat, and the meteoric agencies, these being the ''adaptations'' of the natural theologian.' The author apparently believes that organisation progresses by sudden leaps, but that the effects produced by the conditions of life are gradual. He argues with much force on general grounds that species are not immutable productions. But I cannot see how the two supposed 'impulses' account in a scientific sense for the numerous and beautiful co-adaptations which we see throughout nature; I cannot see that we thus gain any insight how, for instance, a woodpecker has become adapted to its peculiar habits of Life. The work, from its powerful and brilliant style, though displaying in the earlier editions little accurate knowledge and a great want of scientific caution, immediately had a very wide circulation. In my opinion it has done excellent service in this country in calling attention to the subject, in removing prejudice, and in thus preparing the ground for the reception of analogous views.
In 1846 the veteran geologist N. J. d'Omalius d'Halloy published in an excellent though short paper ("Bulletins de l'Acad. Roy Bruxelles,' tom. xiii. p. 581) his opinion that it is more probable that new species have been produced by descent with modification than that they have been separately created: the author first promulgated this opinion in 1831.
Professor Owen, in 1849 ('Nature of Limbs,' p. 86), wrote as follows:- "The archetypal idea was manifested in the flesh under diverse such modifications, upon this planet, long prior to the existence of those animal species that actually exemplify it. To what natural laws or secondary causes the orderly succession and progression of such organic phenomena may have been committed, we, as yet, are ignorant.' In his Address to the British Association, in 1858, he speaks (p. li.) of "the axiom of the continuous operation of creative power, or of the ordained becoming of living things.' Farther on (p. xc.), after referring to geographical distribution, he adds, 'These phenomena shake our confidence in the conclusion that the Apteryx of New Zealand and the Red Grouse of England were distinct creations in and for those islands respectively. Always, also, it may be well to bear in mind that by the word ''creation'' the zoologist means '"a process he knows not what.'' He amplifies this idea by adding that when such cases as that of the Red Grouse are enumerated by the zoologists as evidence of distinct creation of the bird in and for such islands, he chiefly expresses that he knows not how the Red Grouse came to be there, and there exclusively; signifying also, by this mode of expressing such ignorance, his belief that both the bird and the islands owed their origin to a great first Creative Cause.' If we interpret these sentences given in the same Address, one by the other, it appears that this eminent philosopher felt in 1858 his confidence shaken that the Apteryx and the Red Grouse first appeared in their respective homes, 'he knew not how,' or by some process 'he knew not what.'
This Address was delivered after the papers by Mr Wallace and myself on the Origin of Species, presently to be referred to, had been read before the Linnean Society. When the first edition of this work was published, I was so completely deceived, as were many others, by such expressions as 'the continuous operation of creative power,' that I included Professor Owen with other palaeontologists as being firmly convinced of the immutability of species; but it appears ('Anat. of Vertebrates,' vol. iii. p. 796) that this was on my part a preposterous error. In the last edition of this work I inferred, and the inference still seems to me perfectly just, from a passage beginning with the words 'no doubt the type-form,' &c. (Ibid. vol. i. p. xxxv.), that Professor Owen admitted that natural selection may have done something in the formation of a new species; but this it appears (Ibid. vol. nl. p. 798) is inaccurate and without evidence. I also gave some extracts from a correspondence between Professor Owen and the Editor of the 'London Review,' from which it appeared manifest to the Editor as well as to myself, that Professor Owen claimed to have promulgated the theory of natural selection before I had done so; and I expressed my surprise and satisfaction at this announcement; but as far as it is possible to understand certain recently published passages (Ibid. vol. iii. p. 798) I have either partially or wholly again fallen into error. It is consolatory to me that others find Professor Owen's controversial writings as difficult to understand and to reconcile with each other, as I do. As far as the mere enunciation of the principle of natural selection is concerned, it is quite immaterial whether or not Professor Owen preceded me, for both of us, as shown in this historical sketch, were long ago preceded by Dr Wells and Mr Matthews.
M. Isidore Geoffroy Saint-Hilaire, in his lectures delivered in 1850 (of which a Résumé appeared in the 'Revue et Nag. de Zoolog.,' Jan. 1851), briefly gives his reason for believing that specific characters "sont fixés, pour chaque espèce, tant qu'elle se perpétue au milieu des mèmes circonstances: ils se modifient, si les circonstances ambiantes viennent à changer.' 'En résumé, l'observation des animaux sauvages démontre déjà la variabilité limité des espèces. Les expériences sur les animaux sauvages devenus domestiques, et sur les animaux domestiques redevenus sauvages, la démontrent plus clairement encore. Ces memes expériences prouvent, de plus, que les différences produites peuvent etre de valeur générique.' In his 'Hist. Nat. Généralé (tom. ii. p. 430, 1859) he amplifies analogous conclusions.
From a circular lately issued it appears that Dr Freke, in 1851 ("Dublin Medical Press,' p. 322), propounded the doctrine that all organic beings have descended from one primordial form. His grounds of belief and treatment of the subject are wholly different from mine; but as Dr Freke has now (1861) published his Essay on the 'Origin of Species by means of Organic Affinity,' the difficult attempt to give any idea of his views would be superfluous on my part.
Mr Herbert Spencer, in an Essay (originally published in the 'Leader,' March, 1852, and republished in his 'Essays,' in 1858), has contrasted the theories of the Creation and the Development of organic beings with remarkable skill and force. He argues from the analogy of domestic productions, from the changes which the embryos of many species undergo, from the difficulty of distinguishing species and varieties, and from the principle of general gradation, that species have been modified; and he attributes the modification to the change of circumstances. The author (1855) has also treated psychology on the principle of the necessary acquirement of each mental power and capacity by gradation.
In 1852 M. Naudin, a distinguished botanist, expressly stated, in an admirable paper on the Origin of Species ('Revue Horticole, p. 102; since partly republished in the 'Nouvelles Archives du Muséum,' tom. i. p. 171), his belief that species are formed in an analogous manner as varieties are under cultivation; and the latter process he attributes to man's power of selection. But he does not show how selection acts under nature. He believes, like Dean Herbert, that species, when nascent, were more plastic than at present. He lays weight on what he calls the principle of finality, 'puissance mystérieuse, indéterminée; fatalité pour les uns; pour les autres volonté providentielle, dont l'action incessante sur les ètres vivants détermine, à toutes les époques de l'existence du monde, la forme, le volume, et la durée de chacun d'eux, en raison de sa destinée dans l'ordre de choses dont il fait partie. C'est cette puissance qui harmonise chaque membre à l'ensemble, en l'appropriant à la fonction qu'il doit remplir dans l'organisme général de la nature, fonction qui est pour lui sa raison d'ètre.'*
* From references in Bronn's 'Untersuchungen über die Entwickenlungs-Gesetze,' it appears that the celebrated botanist and palaeontologist Unger published, in 1852, his belief that species undergo development and modification. Dalton, likewise, in Pander and Dalton's work on Fossil Sloths, expressed, in 1821 a similar belief. Similar views have, as is well known, been maintained by Oken in his mystical 'Natur-philosophie.' From other references in Godron's work 'Sur l'Espéce,' it seems that Bory St Vincent, Burdach, Poiret, and Fries, have all admitted that new species are continually being produced.
In 1853 a celebrated geologist, Count Keyserling ("Bulletin de la Soc. Gèolog.,' 2nd Ser., tom. x. p. 357), suggested that as new diseases, supposed to have been caused by some miasma, have arisen and spread over the world, so at certain periods the germs of existing species may have been chemically affected by circumambient molecules of a particular nature, and thus have given rise to new forms.
In this same year, 1853, Dr Schaaffhausen published an excellent pamphlet ('Verhand. des Naturhist. Vereins der preuss. Rheinlands,' &c.), in which he maintains the development of organic forms on the earth. He infers that many species have kept true for long periods, whereas a few have become modified. The distinction of species he explains by the destruction of intermediate graduated forms. 'Thus living plants and animals are not separated from the extinct by new creations, but are to be regarded as their descendants through continued reproduction.'
I may add, that of the thirty-four authors named in this Historical Sketch, who believe in the modification of species, or at least disbelieve in separate acts of creation, twenty-seven have written on special branches of natural history or geology.
A well-known French botanist, M. Lecoq, writes in 1854 ('Etudes sur Géograph. Bot.,' tom. i. p. 250), 'On voit que nos recherches sur la fixité ou la variation de l'espèce, nous conduisent directement aux idées émises, par deux hommes justement célèbres, Geoffroy Saint-Hilaire et Goethe.' Some other passages scattered through M. Lecoq's large work, make it a little doubtful how far he extends his views on the modification of species.
The 'Philosophy of Creation' has been treated in a masterly manner by the Rev. Baden Powell, in his "Essays on the Unity of Worlds,' 1855. Nothing can be more striking than the manner in which he shows that the introduction of new species is "a regular, not a casual phenomenon,' or, as Sir John Herschel expresses it, 'a natural in contradistinction to a miraculous, process.'
The third volume of the "Journal of the Linnean Society' contains papers, read July 1st, 1858, by Mr Wallace and myself, in which, as stated in the introductory remarks to this volume, the theory of Natural Selection is promulgated by Mr Wallace with admirable force and clearness.
Von Baer, towards whom all zoologists feel so profound a respect, expressed about the year 1859 (see Prof. Rudolph Wagner, a "Zoologisch-Anthropologische Untersuchungen,' 1861, s. 51) his conviction, chiefly grounded on the laws of geographical distribution, that forms now perfectly distinct have descended from a single parent-form.
In June, 1859, Professor Huxley gave a lecture before the Royal Institution on the 'Persistent Types of Animal Life.' Referring to such cases, he remarks, "It is difficult to comprehend the meaning of such facts as these, if we suppose that each species of animal and plant, or each great type of organisation, was formed and placed upon the surface of the globe at long intervals by a distinct act of creative power; and it is well to recollect that such an assumption is as unsupported by tradition or revelation as it is opposed to the general analogy of nature. If, on the other hand, we view 'Persistent Types' in relation to that hypothesis which supposes the species living at any time to be the result of the gradual modification of pre-existing species a hypothesis which, though unproven, and sadly damaged by some of its supporters, is yet the only one to which physiology lends any countenance; their existence would seem to show that the amount of modification which living beings have undergone during geological time is but very small in relation to the whole series of changes which they have suffered.'
In December, 1859, Dr Hooker published his 'Introduction to the Australian Flora.' In the first part of this great work he admits the truth of the descent and modification of species, and supports this doctrine by many original observations.

 It has been seen in the last chapter that amongst organic beings in a state of nature there is some individual variability; indeed I am not aware that this has ever been disputed. It is immaterial for us whether a multitude of doubtful forms be called species or sub-species or varieties; what rank, for instance, the two or three hundred doubtful forms of British plants are entitled to hold, if the existence of any well-marked varieties be admitted. But the mere existence of individual variability and of some few well-marked varieties, though necessary as the foundation for the work, helps us but little in understanding how species arise in nature. How have all those exquisite adaptations of one part of the organisation to another part, and to the conditions of life, and of one distinct organic being to another being, been perfected? We see these beautiful co-adaptations most plainly in the woodpecker and missletoe; and only a little less plainly in the humblest parasite which clings to the hairs of a quadruped or feathers of a bird; in the structure of the beetle which dives through the water; in the plumed seed which is wafted by the gentlest breeze; in short, we see beautiful adaptations everywhere and in every part of the organic world.
Again, it may be asked, how is it that varieties, which I have called incipient species, become ultimately converted into good and distinct species, which in most cases obviously differ from each other far more than do the varieties of the same species? How do those groups of species, which constitute what are called distinct genera, and which differ from each other more than do the species of the same genus, arise? All these results, as we shall more fully see in the next chapter, follow inevitably from the struggle for life. Owing to this struggle for life, any variation, however slight and from whatever cause proceeding, if it be in any degree profitable to an individual of any species, in its infinitely complex relations to other organic beings and to external nature, will tend to the preservation of that individual, and will generally be inherited by its offspring. The offspring, also, will thus have a better chance of surviving, for, of the many individuals of any species which are periodically born, but a small number can survive. I have called this principle, by which each slight variation, if useful, is preserved, by the term of Natural Selection, in order to mark its relation to man's power of selection. We have seen that man by selection can certainly produce great results, and can adapt organic beings to his own uses, through the accumulation of slight but useful variations, given to him by the hand of Nature. But Natural Selection, as we shall hereafter see, is a power incessantly ready for action, and is as immeasurably superior to man's feeble efforts, as the works of Nature are to those of Art.
We will now discuss in a little more detail the struggle for existence. In my future work this subject shall be treated, as it well deserves, at much greater length. The elder De Candolle and Lyell have largely and philosophically shown that all organic beings are exposed to severe competition. In regard to plants, no one has treated this subject with more spirit and ability than W. Herbert, Dean of Manchester, evidently the result of his great horticultural knowledge. Nothing is easier than to admit in words the truth of the universal struggle for life, or more difficult at least I have found it so than constantly to bear this conclusion in mind. Yet unless it be thoroughly engrained in the mind, I am convinced that the whole economy of nature, with every fact on distribution, rarity, abundance, extinction, and variation, will be dimly seen or quite misunderstood. We behold the face of nature bright with gladness, we often see superabundance of food; we do not see, or we forget, that the birds which are idly singing round us mostly live on insects or seeds, and are thus constantly destroying life; or we forget how largely these songsters, or their eggs, or their nestlings are destroyed by birds and beasts of prey; we do not always bear in mind, that though food may be now superabundant, it is not so at all seasons of each recurring year.
I should premise that I use the term Struggle for Existence in a large and metaphorical sense, including dependence of one being on another, and including (which is more important) not only the life of the individual, but success in leaving progeny. Two canine animals in a time of dearth, may be truly said to struggle with each other which shall get food and live. But a plant on the edge of a desert is said to struggle for life against the drought, though more properly it should be said to be dependent on the moisture. A plant which annually produces a thousand seeds, of which on an average only one comes to maturity, may be more truly said to struggle with the plants of the same and other kinds which already clothe the ground. The missletoe is dependent on the apple and a few other trees, but can only in a far-fetched sense be said to struggle with these trees, for if too many of these parasites grow on the same tree, it will languish and die. But several seedling missletoes, growing close together on the same branch, may more truly be said to struggle with each other. As the missletoe is disseminated by birds, its existence depends on birds; and it may metaphorically be said to struggle with other fruit-bearing plants, in order to tempt birds to devour and thus disseminate its seeds rather than those of other plants. In these several senses, which pass into each other, I use for convenience sake the general term of struggle for existence.